A Drosophila geneticist has identified a strain of flies with a large inversion in the left arm of chromosome 3. This inversion includes two mutations, e (ebony body) and cd (cardinal eyes), and is flanked by two other mutations, sr (stripe thorax) on the right and ro (rough eyes) on the left. The geneticist wishes to replace the e and cd mutations inside the inversion with their wild type alleles; he plans to accomplish this by recombining the multiply mutant, inverted chromosome with a wild-type, inversion-free chromosome. What event is the geneticist counting on to achieve his objective? Explain.