In the vascular bundles of flowering plants, protoxylem is closest to the center of the stem, and

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In the vascular bundles of flowering plants, protoxylem is closest to the center of the stem, and metaxylem is farther out. Is this an endarch or an exarch arrangement? Seed plants always have just this one arrangement, but what about the early vascular plants. Were they endarch or exarch, or did both types occur originally?

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