Males in a certain species of deer have two nonhomologous X chromosomes, denoted X₁ and X₂, and a Y chromosome. Each X chromosome is about half as large as the Y chromosome, and its centromere is located near one of the ends; the centromere of the Y chromosome is located in the middle. Females in this species have two copies of each of the X chromosomes and lack a Y chromosome. How would you predict the X and Y chromosomes to pair and disjoin during spermatogenesis to produce equal numbers of male- and female-determining sperm?