Analysis of unordered yeast tetrads from the cross +++ × a b c yielded the following data:

Which of these genes are linked, and what is the map distance between them?

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Tetrad Number of Class Spores Asci ab c a bc +++ 36 1. +++ a+ c + b+ +++ 2. ab c 14 a++ a++ +b c + bc 3. 32 a++ + b c ++ c 4. 16 5. ab + a b+ ++ c ++ c 2.