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A 34-year-old woman with a history of recurrent urinary tract infections comes to the physician with dysuria and increased urinary frequency. Her urine culture grows colonies of Gram-negative bacteria. The bacteria are isolated and placed in a growth-enhancing nutrient solution, where they undergo rapid cellular division. As they are actively dividing, the bacterial cells are lysed and their DNA is extracted and purified. Analysis of the partially replicated DNA fragments shows the presence of uracil. This finding is most likely mediated by which of the following enzymes