Question: When subpopulations within a metapopulation are highly linked, this can impact the metapopulation in a way not predicted by the Levins metapopulation model, by violating

When subpopulations within a metapopulation are highly linked, this can impact the metapopulation in a way not predicted by the Levins metapopulation model, by violating the assumption that among subpopulations:
colonization rates are equal.
extinction is a random process.
interpatch distances are not equal.
geometric growth rates are equal.
patch sizes are equal.
When subpopulations within a metapopulation are

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