DNA can be modeled as an elastic rod which can be twisted or bent. Suppose a DNA molecule of length L is bent such that it lies on the arc of a circle of radius R_c. The reversible work involved in bending DNA without twisting is w _bend =BL/2R²_c where B is the bending force constant. The DNA in a nucleosome particle is about 680. Å in length. Nucleosomal DNA is bent around a protein complex called the histone octamer into a circle of radius 55 Å. Calculate the reversible work involved in bending the DNA around the histone octamer if the force constant B = 2.00 × 10⁻²⁸ J m−1.