Sometimes Billy does cuddle with Kathy even if he doesn't feel very affectionate himself, but it's not a very enjoyable experience for him. Which proposition of Affection Exchange Theory explains what Billy is experiencing?

A) The need and capacity for affection are inborn

B) Affectionate feelings and affectionate expressions are distinct experiences that often, but need not, covary

C) Affectionate communication is adaptive with respect to human viability and fertility

D) Humans vary in their optimal tolerances for affection and affectionate behavior

E) Affectionate behaviors that violate the range of optimal tolerance are physiologically aversive

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